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The people of the Hoan G'Tay Sho "gifted" members of their tribe (wise ones, old ones, young ones...) regularly to the "Spirits Who Dwell On High" and Kakuk lived most of his life as a slave in Winnowill's personal chambers.
When he and the other humans were ordered by Winnowill to kill the WolfRiders in their pursuit to reclaim Suntop, who she'd kidnapped, the humans were much relieved when the Wolfriders took to the rafters where they couldn't follow as they didn't want to have their "gods" blood on their hands.